Thursday, 2 April 2026

Signal Without Semiosis: Value, Selection, and the Misreading of Meaning in Biology — 3 The Wallace Temptation

If Darwin’s account of sexual selection can proceed without invoking meaning, why does the language of signalling return so persistently?

Why do biological traits so readily come to be described as if they represent something?

To answer this, we can turn—carefully—to a contrasting tendency associated with Alfred Russel Wallace.

The point here is not historical attribution in a strict sense, but conceptual orientation.

What matters is the temptation.


From preference to indication

Where Darwin begins with preference, an alternative move begins with indication.

Instead of asking:

which traits are taken up within a system of selection?

the question shifts to:

what do these traits reveal?

In this framing:

  • elaborate features are not just selected

  • they are taken to indicate underlying qualities

  • strength, health, or genetic fitness

The tail does not simply participate in selection.

It becomes evidence.


From indication to representation

Once a trait is treated as indicating something, a further step is close at hand.

If a feature reliably indicates an underlying condition, then:

it can be treated as standing for that condition.

At this point, the language of signalling becomes almost inevitable:

  • the tail signals fitness

  • the display communicates quality

  • the behaviour conveys information

The system is now described in semiotic terms.


The structure of the temptation

The movement can be summarised as a sequence:

  1. Correlation
    A trait co-occurs with some condition.

  2. Indication
    The trait is taken as evidence of that condition.

  3. Representation
    The trait is treated as standing for that condition.

  4. Signal
    The system is described as one of communication.

Each step appears reasonable.
Together, they produce a conceptual shift.


Where the shift occurs

The crucial transition is from:

indication → representation

Indication is a relation we, as observers, can draw:

  • we notice that one feature tracks another

  • we treat it as informative

But representation requires something more:

  • a system in which one form functions as another

  • a relation internal to the system, not just observed from outside

The temptation lies in moving from one to the other without marking the difference.


The observer’s projection

Part of the difficulty is that the relation between trait and condition is often visible to us.

We can measure:

  • correlations between ornament and health

  • statistical associations between display and reproductive success

From this vantage point, it is natural to describe the trait as informative.

But this is an observer’s description.

It does not follow that:

  • the organism treats the trait as a representation

  • or that the system operates semiotically

We may be projecting our own interpretive framework onto the system.


Reliability is not meaning

The temptation is strengthened by reliability.

If a trait:

  • consistently correlates with a condition

  • and consistently elicits a response

then it begins to look like a signal.

But reliability alone does not establish semiosis.

A system can be:

  • reliably responsive

  • without being meaning-making

Consistency of outcome does not imply the presence of representation.


The drift into communication

Once representation is assumed, the rest follows easily.

  • representation implies content

  • content implies transmission

  • transmission implies communication

And so the biological system is redescribed as one in which:

information is exchanged between organisms.

At this point, the original value-based dynamics of selection are no longer foregrounded. They are reframed as instances of communication.


What is lost

In this shift, something important is obscured.

The explanatory power of:

  • differential uptake

  • preference

  • and selection

is replaced by an appeal to:

  • signalling

  • information

  • and interpretation

But unless semiosis is independently established, this move adds description without adding explanation.


The persistence of the temptation

The Wallace temptation persists because it aligns with powerful intuitions:

  • that behaviour is guided by information

  • that organisms respond to what things mean

  • that coordination requires communication

These intuitions are not groundless.
But they may not apply in the way they are assumed to.


Reframing the issue

We can now restate the problem more precisely:

when we describe a biological trait as a signal, are we identifying a semiotic relation, or reinterpreting a value-based process in semiotic terms?

The distinction matters.

Because if the latter is the case, then:

  • meaning is being inferred where it is not required

  • and the concept of the signal is doing more conceptual work than it can support


Transition

Having identified the temptation, the next step is to return to the underlying dynamics.

If meaning is not required to explain these systems, what is?

What kind of process is actually doing the work of coordination, selection, and stabilisation?

To answer this, we return to the notion introduced earlier:

value as a non-semiotic dynamic.

It is here that the explanatory weight lies.

No comments:

Post a Comment